[Title Page] [List of Genera] Comparison of Drawings] [Regional Keys to Genera] [Descriptions of New Taxa] [Geographical Distribution] [Color Photographs] [Scanning Microphotographs] See also Catalogue of Salticidae [ Title page] List of Genera - names beginning with: A BC DEFG HIJKL MN OPQ RS TUVWXYZ References: Authors beginning with: A BC DEFG HIJKL MN OPQ RS TUVWXYZ[Key for Salticidae of Levant][Simon's 1899-1902 Keys to the Salticid Groups]
by Jerzy Proszynski 1997
Keys for Identification of Salticidae
Salticidae Genera of Central Europe
by Jerzy Proszynski
Salticidae are medium size or small spiders, recognizable by unique anterior end of body broadly truncated, with 2 pairs of eyes on the face (anterior surface): large anterior median (AME), and smaller by half anterior lateral (ALE). Remaining two pairs of eyes (of II and III row) located on dorso-lateral edges of cephalothorax, delimit long quadrangle called eye field (taking about half of cephalothorax); body compact, legs short or slightly longer, move by jumping and running; active hunters with complex behavior. European genera spun silky retreats and cocoons but not catching webs family Salticidae .
1. Ant-like (abdomen constricted, petiolus long,not hidden beneath anterior part of abdomen) - ant like 2 -Non ant-like - 4 2 (1). Medium size spiders - 3 - Very small spider, cephalothorax not constricted, flat and low . FEMALE epigynum sclerotized, smooth, openings located in mid-length, usually somewhat laterally, spermathecae very small; MALE Bulbus small pear shaped with a white membraneous spot corresponding with inflatable receptacle, embolus located anteriorly and short - Synageles
3 (2). Eye field distinctly elevated above thorax, which is constricted; FEMALE epigynum - White membraneous area located posteriorly, divided medially by internal sclerotized channels, prominently visible through membrane, MALES small coil of seminal receptacle in anterior half of round bulbus, embolus making 2 or more loops around the bulbus, tibial apophysis very small and hook like bent - Myrmarachne - Eye field and cephalothorax flat, on the same level: FEMALE Copulatory openings located anteriorly and separated by median ridge stretching over anterior half of epigynum, channels are straight and thickwalled, like spermathecae; MALE Embolus broad, about 1/3rd width of bulbus, arising in mid length of bulbus from its swollen basal part (ant like spider) - Leptorchestes
4(1) Very small species, shorter than 4 mm (except. some Sitticus caricis), (with length of eye field about 40 % in % of length of cephalothorax, cephalothorax low - about 45-60 % 1 at eyes III) - 5 - Medium size or large species, from 4 to 12 mm long - 7
5(4) Dark colored, tegument reflecting light in MALES whole body and abdomen with scutum, seminal receptacle channel meandering, embolus coiled atop bulbus, in some species in a form of a thick fleshy process. FEMALES only cephalothorax is light reflecting but abdomen is soft and dark grey, height of cephalothorax about 45 %; epigynum weakly sclerotized usually with single white membraneous "window", more rarely two - oval or round, spermathecae round or oval, visible as darker spots behind the "window" (Very xerothermic occurence, Northernmost in Czech Rp.) - Chalcoscirtus - Tegument not reflecting light, MALE without abdominal scutum - 6
6 (5) Abdominal pattern with dark chevrons on light background or entirely dark, sometimes also with a few pairs of small light dots, cephalothorax broader than eyes III, eye field about 45 %, cephalothorax slightly higher - 60 %). MALE embolus coiled atop bulbus (sometimes around hollow pit or coiled hidden within that pit), seminal receptacle channel meandering forming broad loops. FEMALE epigynum like in Chalcoscirtus - their separation uncertain, always require preparation of epigynum; (common Euophrys frontalis in forest litter or mosses) - Euophrys ; Pseudeuophrys - Abdominal pattern form a delicate dark network on light background, connected with thin median chevrons, cephalothorax broadest among the three genera - its width at eyes III 76 %, low (45% . MALE Embolus making short coil on the side of bulbus, with smallprotuberance inside covered with spikes (reticulatus, robustus and valentulus). FEMALE epigynum with two grooves surrounded by sclerotized black rims instead of membraneous "windows". N. reticulatus - common species living in forest litter or mosses) - Neon (ATTENTION - rayi DIFFERENT] - MALE with short apical embolus atop long fleshy triangular trunk, tibial apophysis short. FEMALE epigynum weakly sclerotized depression between bent rims, but without posterior rim, arching anteriorly, spermathecae spherical, posterior with long thin channels laterally, openings under anterior rim (Oriental genus penetrating Eastern Palaearctics, P. castriesiana has Pliocene relict populations in S Europe ) - Phintella
7 (4) - Palpal organ and epigynum resembling Euophrys; but MALE with fringe of black setae along tibia and metatarsus III; FEMALE abdomen with scattered, small, dark dots, epigynum with windows extending over the whole length of epigynum (two Mediterranean species Saitis barbipes + tauricus) - Saitis - Palpal organ and epigynum different from Euophrys - 8
8 (7) Cephalothorax broad - 9 - Cephalothorax narrow with sides more or less parallel - 12
9 (8) Eye field relatively long, takes about half of length of cephalothorax, broadest at eyes III, eyes at edges of dorsal surface - 10 - Eye field distinctly shorter than half of cephalothorax, it sides approximatively parallel, narrower than cephalothorax, which is high, somewhat cone shaped, with narrow eye field atop, its basal outline almost circular, usually swollen beneath eyes III. epigynum with white anterior window, narrow and square, coiled membraneous channels. Palpal organ with bulbus round with triangular flap lik protuberance, twice encircled by embolus. (Widespread species, occuring from Oriental Region, Africa to Mediterranean, in Europe only in the South) - Thyene imperialis
10(9) Posterior slope of cephalothorax broad and steep, margin of abdomen extends above posterior part of thorax, particularly striking in MALES, surface of carapace rough, light reflecting, abdomen broader than cephalothorax. Eye field broadening posteriorly and taking longer than half of flat dorsal surface of cephalothorax. MALE embolus making tight transversal coil atop bulbus, but seminal receptacle duct does not make large loops. FEMALE copulatory openings in anterior median part of pit with surrounding rim unsclerotized, channels running medially posteriorwards and making coils in the posterior part of the pit - Ballus - Posterior slope of cephalothorax narrowing and gently sloping, surface of carapace smooth, abdomen almost black, often with with pairs of white spots. MALE bulbus round, slightly flattened apically, or transversally oval, with embolus arising lateraly and making small bent. FEMALE anterior half of epigynae white, membraneous, oval, with prominent vaginal sclerotized roof in the middle - Bianor
11 - Abdominal pattern consists of pairs of transverse marginal whitish spots on darker, reddish or greenish background. MALE bulbus bag shaped with twin sclerotized processes, short, atop white apical pad. FEMALE epigynum copulatory openings located anteriorly, often in a distinct semilunar groove. (Live on pine trees: D. hastatus on young pines, D. rudis on pine crowns) - Dendryphantes - Abdominal pattern consists of pairs of marginal darker spots on whitish background. MALE embolus long, gently bent, arising anterolaterally on anterior bulbus. FEMALE copulatory openings located antero-laterally, but not in a semilunar groove, channels converging posteriorwards, scleroti zation of the epigynum less striking - Macaroeris
12(8) Cephalothorax low and long, legs I strikingly larger than II-IV, particularly in MALES - 13 - Cephalothorax high, if legs I larger then not so strikingly - 15 - Profile of cephalothorax high, body "hairy" - 16 - Cephalothorax giving no clear impression of being either high or low, just "average" - 17
13(12) - A row of stridulatory spines on tubercles under eyes lateral and corresponding microspines prolaterally on femur I, tibia I swollen medially with reduced rudimentary spines or devoid of them, and with several long an thin, usually bent sensory hairs, small spider - Pseudicius - Spider without row of stridulatory spines under eyes lateral, tibia not swollen, armoured with normal spines - 14
14(12) Abdomen particularly long, MALE palpal organ with flattened lateral surface, epigynum without external sculpture, FEMALE epigynum in a form of a smooth plate with only openings, short initial channels pas into komplicated knots of long loops, without distinct spermathecae. (Marpissa radiata lives on shores of waterbodies and watercourses: retreats and cocoons located usually on: bent leaves of Phragmitis, M. muscosa on bark of pine tree trunks) - Marpissa -- Abdomen not only long but also very narrow . FEMALE epigynum in a form of a smooth plate with only single median openings or shallow depressions (Mediterranean, farther North only along sea beaches of Atlantic and North Sea ) - Marpissa nivoyi - FEMALE epigynum in a form of a smooth plate, with copulatory openings hidden under posterior edge of epigynum, that edge is carved into two bays separating a mendian "promontory" in a form of a small triangle, short cannals run from the sides of triangle anteriorwards and turns into short, coiled sclerotized spermathecae, translucent by tegument as dark spots (Mediterranean, reported also from reeds on shores of Neusiedlersee in Austria ) - Mendoza canestrini - Body not as long 15
15(12) Cephalothorax usually with longitudinal light and dark stripped pattern along thorax and on abdomen, copulatory organs in MALE of "Aelurillus" type (bulbus relatively large oval, flattened - at least apically, embolus hidden beneath it with only tip protruding from under anterior edge, palpal tibia apophysis always bifurcated, rami separated by "U" shape grove. In FEMALES posterior openings (large and prominent located posteriorly, either medially or near posterolateral angle of epigynum) and coiled spermathecae, medial, with internal convoluted chambers. Living on sand. - Phlegra - Similar to Phlegra, but body uniformly dark, without stripes or chevron mark on the eye field. MALE ventral apophysis narrow and slightly bent laterally, FEMALE epigynum with openings near posterior corners, copulatory channel running straight, then bent, spermathecae convoluted median.(On sunny, warm rocky slopes) - Asianellus - Cephalothorax apears higher than in Phlegra, no striped pattern (in MALES usually with whitish or silver chevron mark on the eye field); palpal organ with ventral tibial apophysis triangular, separated from the dorsal apophysis by "V" shape grove. FEMALES: epigynum with two "wings" - sclerotized diagonal edges, spermathecae in a form of complicated sclerotized chambers. On warm sandy grounds or sunny slopes - Aelurillus
16(12) Profile of cephalothorax triangular, with eye field sloping forward, eyes III area forming the highest part and posterior slope begining right beyond the eye field, tarsi I-II with scopulae, retrolateral margin of chelicerae and tooth not developped, prolateral one with a thin sclerotized edge ending with a small tooth pointed forward. FEMALE epigynum highly sclerotized with openings located anteriorly, median part elevated, triangular posterior rim pushed inside abdomen. MALE pedipalpal femur halfcrescent and broad, often with small ventral protuberance, embolus always folowed by conductor.(Living on sand, cryptically colored) - Yllenus - Profile of cephalothorax rounded, colouration dull greyish with ill defined lighter spots, palpal organ and internal structure of epigynum characteristic - Carrhotus - FEMALES and immature MALES black with two long white spots on abdomen, sometimes two regular parallel white lines; in MALES abdomen red with longitudinal black streak encircled by a thin white margin.Very common in Mediteranean, in C Europe on isolated xerothermic spots. FEMALE Only roofs of copulatory openings are sclerotized and form sharp triangle in the anterior part of epigynum, followed by broad channels running posteriorwards, remaining part of epigynum not sclerotized and covered by long dense setae MALE Embolus very long, posterior half of bulbus makes long narrow protuberance bent under angle - Philaeus - Dark median serrated abdominal streak on light narrow background margins and sides dark; lateral protuberance on bulbus, embolus arising in the 5 o'clock position, encircling 3/4th of bulbus, tibial apophysis long, diagonal indistinctly broadened terminally; epigynum oval with bent slits, broad membraneous channels and small sclerotized spermatheca (Mediterranean species, with numerous relatives in Mediterranean, S Palaearctics as well as arid Africa, Near East and Oriental Region) - Mogrus
17(12) - Abdomen uniformly colored, with or without white pattern - 18 - Abdomen with irregular dark lines, or mosaic of dark, brown or grey spots and patches of various sizes, with or without white pattern - 19 - Abdomen with broad median light longitudinal streak, posteriorly with a pair of lateral spots and a few chevrons, extended anteriorly by light thoracal streak, in some forms reaching eyes I; bulbus expanded laterally with short bent embolus anteriorly; epigynum sclerotized with posterior median groove; large or medium size spider (Widespread, pantropical genus, one species occuring also in the Mediterranean [but several in Israel], locally in S Europe, in C Europe in hot houses) - Plexippus - Striking bright coloration (particularly in MALE): cephalothorax and abdomen anteriorly pink, posteriorly black (fig.); palpal organ with anterior embolus running around anaterior and lateral edge of bulbus (typical for Spartaeinae) and long articulating tibial apophysis , epigynum characteristic shield with sclerotized edge, spherical spermathecae and anteriorl arches of channels ( Mediterranean species, related Oriental species penetrating S Palaearctics) - Cyrba algerina
18(17) Abdomen uniformly black irridescent blue, violet or green, light reflecting due to colourless scales, in some species one or more pairs of small white marginal spots and a semicrescent anterior line, legs often yellow, medium - small spiders. MALE pedipalpal femur with large horn like protuberance (single, bifurcated or trifurcated). FEMALE epigynum with either single or twoseparate sclerotized depression round or transversally oval, sometimes partially or entirely surrounded by an elevated rim, copulatory openings usually located laterally or, more rarely posteromarginally, or hidden under rim, with channels and spermathecae short and simple - Heliophanus - Abdomen dark with pairs of transversal or diagonal white stripes or their rudiments ("zebra spider") but no median line, in MALES chelicerae overgrown. MALE palpal organ with large and broad apophysis, bag shaped bulbus and short embolus. FEMALE epigynal depression extended by posterior elongate part, sometimes anteriorly hidden under a roof like rim, channels and spermathecae more complicated than in Heliophanus - Salticus - Abdomen black with contrasting white line, continuous or interrupted, in some species there are one or two pairs of diagonal white lines marginally and thick anterior line, on cephalothorax in some species white spots behind eyes III and median white line. MALE Bulbus oval with embolus usually bifurcated, tibial apophysis supported on some kind of swelling of cymbium (in P. tripunctatus three swellings, apophysis is grown into wall of cymbium) FEMALE Entire epigynum sclerotized with median ridge separating two semicircular grooves, location of vaginal opening varies from anterior to posterior in various species - Pellenes
19(17) Abdomen with pairs of dark diagonal lines, usually on darkish or even entirely black background giving spiders dull blackish appearance, sometimes with white marginal lines on abdomen and sides of cephalothorax. Relatively large spider with characteristic genital organs. MALE Bulbus large - round, oval or narrowing posteriorly, embolus relatively broad, protruding from under anterior median angle of bulbus, apophysis of medium length or longer. FEMALE anterior half of epigynum white membraneous, the posterior being strongly sclerotized (in some speces with postero-laterl sclerotized pockets) in E. laetebunda reduced to narrow posterior rim - Evarcha - Abdomen with mosaic dark, brown or grey spots and patches of various sizes, usually with a pair of median white spots slightly behind mid-length of abdomen, as well as some chevrons - 20 20(19) Genital organs of two related types, very characteristic cheliceral dentition: retrolateral margin of chelicerae not developed, a tooth with several conical cusps on prolateral margin. In a few species no white spots, males of some high mountain species have abdomen black, bissected by white longitudinal line - in Europe only Italian Alpine Sitticus longipes] a) FEMALE epigynum with a central pit under whose sclerotized rim copulatory openings are hiden, channels running outwards from the pit. MALE bulbus round, ventrally flattened, with embolus arising laterally or basally and making either half circle arround bulbus or encircling it once or twice, with characteristic loop of seminal receptacle canal, tibial apophysis small or very large. Sitticus floricola and related b) FEMALE epigynum with copulatory openings in a form of spaced diagonal slits in the mid length of surface of epigynum, spermathecae form single bends in the posterior half of epigynum. MALE - bulbus elongate oval or bag shaped, embolus slightly longer than bulbus arising usually at its latero basal angle, small tibial apophysis - Sitticus distinguendus and related c) FEMALE epigynum in a form of simple depression, followed posteriorly by two less sclerotized lobes, spermathecae - centrally located sclerotized simple chambers without channels. MALE bulbus round, embolus short and thick arising laterally, apophysis large - Sitticus pubescens - Pairs of white abdominal spots larger, single conical retrolateral cheliceral tooth, genital organs dif ferent, in MALE cymbium dorsally with large white spot( numerous Mediterranean species only) - Habrocestum - Male pedipalpal tibia long, with long mane of white setae, genical organs of different type; single bicusps retrolateral cheliceral tooth, (cosmopolitic species in warm countries, in C Europe in greenhouses). - Hasarius ================================================================== This English Language version of the original Key, written by myself, was a basis for abbreviated and modified version published in German Language as a chapter "Salticidae Springspinnen" by J. Proszynski, in edited by S. Heimer and W. Nentwig: "Spinnen Mitteleuropas", Verlag Paul Parey, 1991. The Editors omitted my name at the heading of the chapter. Made accessible for Internet with permission of Verlag Paul Parey.
Copyright © for the page by J. Proszynski , 2000.